A Rarely Seen Taxonomic Revision with Immense Value for 41 Years and Counting: Reflections on the 1981 Monograph of Trichonta Winnertz, 1864 (Diptera: Mycetophilidae) by Raymond Gagné, with an Integrative Revision of the Trichonta Vulcani (Dziedzicki, 1889) Species Complex

We celebrate Raymond J. Gagné for his contributions to taxonomy of the Mycetophilidae (Diptera), specifically forty-one-years-old monograph Holarctic Trichonta Winnertz, 1864 that is still primary source used species identification in genus. briefly reflect on monograph's impact and demonstrate by use recent DNA barcode data extracted from BOLD Systems (BOLD) model distribution presented holds up scrutiny. To refined concept now being applied an integrative taxonomic approach includes barcodes, we revise a small, but distinct, complex recognized as one morphologically defined example old pan-Holarctic taxon, vulcani (Dziedzicki, 1889). find (sensu Kallweit 1998) consist at least six Region which three are described new science: japonica Kurina, (East Palearctic), neovulcani Kjaerandsen, Nearctic), raymondgagnei (Holarctic), trifida Lundstrom, 1909 (wide tristis (Strobl, 1898) 1889) Palearctic). All distinctly separated barcodes correspond well minor, constant, differences their male terminalia. However, widespread species, trifida, displays some genetic morphological differentiation between western eastern Palaearctic populations. presently consider these populations conspecific pending broader sampling. further propose replacement name nepalensis superba Gagné, 1981, junior homonym Ostroverkhova, 1979, latter secondary synonym 1898).